īased on the size of the vertebrae, Titanoboa is the largest snake in the paleontological record. The teeth themselves are weakly ankylosed, meaning they are not strongly connected to the jawbone. According to it, Titanoboa is unique in the high amount of palatal and marginal tooth positions compared to others boids, the quadrate bone is oriented at a low angle and the articulation of both the palatine to pterygoid and pterygoid to quadrate are heavily reduced. The skull is only briefly described in a 2013 abstract. They are robust with a uniquely T-shaped neural spine. Most material of Titanoboa consists of vertebrae that in life would be located before the cloaca. The relative size of Titanoboa to the modern human, Gigantophis, reticulated python, and green anaconda. The species name on the other hand is a reference to the Cerrejón region. The scientific name combines the Greek word "Titan" with Boa, the type genus of the family Boidae. Field work continued following these initial discoveries, recovering multiple additional specimens including three skulls with associated postcranial bones. The snake was discovered on an expedition by a team of international scientists led by Jonathan Bloch, a University of Florida vertebrate paleontologist, and Carlos Jaramillo, a paleobotanist from the Smithsonian Tropical Research Institute in Panama. Before this discovery, few fossils of Paleocene-epoch vertebrates had been found in ancient tropical environments of South America. These specimens consist of the holotype, a large precloacal vertebrae, the paratype, also a vertebrae, and 184 additional remains identified as additional vertebrae and ribs (some of which found in articulation), amounting to a total of 28 specimens in addition to the holotype and paratype. cerrejonensis were found in the Cerrejón Formation of the coal mines of Cerrejón in La Guajira, Colombia. In 2009, the fossils of 30 individuals of T. Main articles: Cerrejón Formation and Cerrejón
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